Entangled Life by Gillian Barker Eric Desjardins & Trevor Pearce

Author:Gillian Barker, Eric Desjardins & Trevor Pearce
Language: eng
Format: epub
Publisher: Springer Netherlands, Dordrecht

4 Type Fitness and Coarse-Grained Environmental Variation

The preceding section provided a framework for thinking about how fitness depends on environmental variation. In this section, I’ll discuss in more detail my claim that parts of the environment with a significant probability of recurring—of being experienced repeatedly by members of a population, over generational time—give rise to fitness in a sense different than do circumstances that are idiosyncratic to a particular time, place, configuration of organisms, etc. I begin by relating Brandon’s previous discussion of environmental variation to my view (§4.1). I then discuss the how one might draw a formal distinction between circumstances and other subenvironments, thus placing a lower limit on what kinds of subenvironments are worth modeling or studying empirically (§4.2). I explain how token fitnesses and environmental circumstances might contribute to type fitnesses relative to larger subenvironments (§4.3), and respond to a challenge that environmental gradients pose for my view (§4.4).

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